rhizomes mostly long-creeping, often siphonostelic or polystelic, bearing jointed hairs; petioles often with epipetiolar buds, usually with a gutter-shaped vascular strand (open end adaxial); blades often large, 2–3-pinnate or more divided; indument of hairs; veins free, forked or pinnate, rarely anastomosing and then without included veinlets; sori marginal or submarginal, linear or discrete, indusia linear or cup-like at blade margins, or reflexed over sori; spores tetrahedral and trilete, or reniform and monolete; gametophytes green, cordate; x = 26, 29, 30, 31, 33, 34, 38, 46, 47, 48, and probably others.
